My primary academic interests are in evolution, dispersal ecology, and plant-animal interactions. My research has used diverse groups of animals, including forest herbs, ants, bats, bluebirds, and fish of headwater streams. I helped to start the Brown Family Environmental Center and serve as its Executive Director. My eduction includes an A.B. from Kenyon College and Ph.D. from Stanford University. I taught at Northwestern University before coming back to Kenyon in 1980.
Ecology of mutualism, evolution of sex ratios, emergent behaviors of social insects
1973 — Doctor of Philosophy from Stanford University
1968 — Bachelor of Arts from Kenyon College